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Wolverine

 

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A North American Wolverine looks at the camera, crouched on a vibrant green and brown forest floor. The wolverine's fur is a chestnut brown (on the head) and dark brown-black
Information icon Photo by Roy Anderson. View on Flickr

Wolverine (Gulo gulo luscus)

Current Status: Not Listed

Species description: The wolverine is the largest terrestrial member of the family Mustelidae.  Adult males weigh 12 to 18 kilograms (26 to 40 pounds) and adult females weigh 8 to 12 kilograms (17 to 26 pounds).  The wolverine resembles a small bear with a bushy tail.  It has a broad, rounded head; short, rounded ears; and small eyes.  Each foot has five toes with curved, semi-retractile claws used for digging and climbing.

Location: In North America, wolverines occur within a wide variety of habitats, primarily boreal forests, tundra, and western mountains throughout Alaska and Canada; however, the southern portion of the range extends into the contiguous United States.

Currently, wolverines are found in the North Cascades in Washington and the Northern Rocky Mountains in Idaho, Montana, Oregon (Wallowa Range), and Wyoming.  Individual wolverines have also moved into historic range in the Sierra Nevada Mountains of California and the Southern Rocky Mountains of Colorado, but have not established breeding populations in these areas.  

Click here to view Wolverine photos on Flickr


Distribution, Breeding, Diet, & Range »

Distribution: Research indicates that wolverines either did not exist as established populations or were extirpated prior to settlement and to the compilation of historical records in the Great Lakes region, possibly due to climate changes that occurred through the 1800s and 1900s.  The widely scattered records from this region are consistent with dispersing individuals from a Canadian population that receded north early in the 1800s.  The possibility that wolverines existed as established populations prior to the onset of trapping in this area cannot be ruled out, but we have no evidence that they did.  No evidence in the historical records suggests that wolverines were ever present as established populations in the Great Plains, Midwest, or Northeast.

The delineation of wolverine historical and present distribution is inherently difficult for several reasons.  Wolverines tend to live in remote and inhospitable places away from human populations.  Wolverines naturally occur at low densities and are rarely and unpredictably encountered where they do occur.  Wolverines often move long distances in short periods of time when dispersing from natal ranges, making it difficult or impossible to distinguish with confidence between occurrence records that represent established populations and those that represent short-term occupancy without the potential for establishment of home ranges and reproduction.  These natural attributes of wolverines make it difficult to determine their present range, or trends in range expansion or contraction that may have occurred in the past.

Breeding: Breeding generally occurs from late spring to early fall.  Females undergo delayed implantation until the following winter to spring, when active gestation lasts from 30 to 40 days.  Litters are born between February and April, containing one to five kits, with an average in North America of between 1 and 2 kits. 

Female wolverines use natal (birthing) dens that are excavated in snow. Persistent, stable snow greater than 1.5 meters (5 feet) deep appears to be a requirement for natal denning, because it provides security for offspring and buffers cold winter temperatures.

Diet: Wolverines are opportunistic feeders and consume a variety of foods depending on availability.  They primarily scavenge carrion, but also prey on small animals and birds, and eat fruits, berries, and insects.  Wolverines have an excellent sense of smell that enables them to find food beneath deep snow.

Range: Wolverines require a lot of space; the availability and distribution of food is likely the primary factor in determining wolverine movements and home range size.  Wolverines travel long distances over rough terrain and deep snow, and adult males generally cover greater distances than females.  Home ranges of wolverines are very large, but vary greatly depending on availability of food, gender, age, and differences in habitat.  These home range sizes are large for mammals of the size of wolverines and may indicate that wolverines occupy a relatively unproductive niche. 

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Recent Actions »

October 8, 2020: The best available science show that the factors affecting wolverine populations are not as significant as believed in 2013 when the U.S. Fish and Wildlife Service (Service) proposed to list the wolverine found in the contiguous United States as threatened.

New research and analysis show that wolverine populations in the American Northwest remain stable, and individuals are moving across the Canadian border in both directions and returning to former territories. The species, therefore, does not meet the definition of threatened or endangered under the Endangered Species Act (ESA). Accordingly, the Service has withdrawn its listing proposal.

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Withdrawal Announcement FAQs »

Q: Why is the U.S. Fish and Wildlife Service announcing this withdrawal today?

A: On February 4, 2013, we published a proposed rule to list a distinct population segment of the North American wolverine found in the contiguous United States as threatened. On August 13, 2014, we withdrew that proposed rule based on our conclusion that the factors identified as affecting the distinct population segment were not as significant as believed at the time of its publication.

In October 2014, three complaints were filed in the District Court for the District of Montana by Defenders of Wildlife, WildEarth Guardians, Center for Biological Diversity and other organizations challenging the withdrawal. Numerous parties intervened in the litigation. These three cases were consolidated, and on April 4, 2016, the court issued a decision. As a result, the August 13, 2014, withdrawal was remanded to the Service for further consideration consistent with the order, returning the process to the proposed rule stage and the status of the wolverine under the Endangered Species Act to that of a proposed species.

Today’s decision completes our response to the 2016 court order and again withdraws our 2013 proposal to list the species. This decision reflects the latest and best available science, including improvements to our understanding of the biology and behavior of the species, as well as advances in snow and climate modeling.

Q: How many wolverines are in the contiguous United States?

A: By their nature, wolverines are difficult animals to survey. Populations occur in highly remote areas and in naturally low densities across their North American range due, in large part, to their need for large, exclusive territories. At present, there is no reliable estimate of the number of wolverines that currently occupy or previously occupied the contiguous United States, nor are there reliable quantitative estimates of wolverine population trends. The often-cited population estimate of 318 wolverines (range: 249–926) is derived from a 2013 model to predict suitable habitat that also estimated the potential wolverine capacity to be 644 individuals. However, these estimates did not consider important spatial considerations related to wolverine behavior, such as territoriality, and are not a definitive population count across the western United States.

As discussed in the wolverine species status assessment, preliminary field results from the state wildlife agencies’ 2016–2017 occupancy study across Idaho, Montana, Washington and Wyoming report a total of 86 photographic detections through camera-trapping and 157 wolverine hair samples collected for genetic analysis. It has not yet been determined from the camera-trap images or hair samples how many of the detections are unique individuals.

There are many fewer wolverines in the contiguous United States than there are in Canada and Alaska, but this is most likely a result of the amount of naturally occurring habitat available within the contiguous United States, both currently and historically, for a species that needs large, exclusive territories, and is not a reflection of poor conservation.

Q: What is a Species Status Assessment?

A: A species status assessment is a focused, repeatable and rigorous scientific assessment of biological risk that aids decision makers who must use the best available scientific information to make policy decisions under the Endangered Species Act. It begins with a compilation of the best available information on the species’ life history, habitat and taxonomy. Next, it describes the current condition of the species’ habitat and demographics (i.e. how the population changes over time), and the probable explanations for past and ongoing changes in abundance and distribution within the species’ range. Last, a species status assessment forecasts the species’ response to probable future scenarios of environmental conditions and conservation efforts.

Q: How long into the foreseeable future did the Service forecast its projection to determine whether the wolverine would meet the definition of “threatened?”

A: The future time frame evaluated in our analysis is approximately 38 to 50 years, which includes the potential for observing several generations of species and captures our best professional judgment of the projected future conditions related to trapping/harvesting, climate change, or other potential cumulative impacts. We note in our species status assessment that evaluations of future conditions for the species have an inherent level of uncertainty relative to demographic risks, particularly those related to climate change projections, and beyond this range of time, modeling uncertainty increases substantially.

Q: Why is the wolverine not considered a distinct population segment? 

A: In light of the updated analysis and new information in the wolverine species status assessment, we re-evaluated whether wolverines in the contiguous United States meet the criteria of a distinct population segment. According to  our policy, a population must be both discrete in relation to the remainder of the species, and significant to the remainder of the taxon to which it belongs in order to qualify. We have concluded that the population of wolverines in the contiguous United States is not discrete, as defined in our policy.

Wolverines currently occupying the contiguous United States do not meet the definition of discrete because they are a metapopulation that is genetically connected to the wolverines in Canada. New information from genetic and observational studies shows that wolverines in the lower 48 are connected to populations in Canada and Alaska, these populations interact on some level, and migration and breeding is possible between groups. For example, wolverines in the North Cascades region of Washington are known to move into British Columbia. Furthermore, there are no differences in management or conservation status between wolverines in the lower 48 and Canada that are significant or represent an inadequacy of existing regulatory mechanisms to conserve the species. Therefore lower 48 wolverines are not discrete from the remainder of the taxon. This means that the wolverines in the lower 48 states do not qualify as a distinct population segment and they are instead an extension of the population of wolverines found further north, a positive sign for the long-term health of the species.

Q: What does the science show in regard to the impact of climate change on wolverines?

A: Climate change model projections for the range of the wolverine within the contiguous United States indicate increases in temperature by the mid-21st century. Snow cover is projected to decline in response to warming temperatures and changing precipitation patterns, but this varies by elevation, topography and geographic region. In general, models indicate higher elevations will retain more snow cover than lower elevations, particularly in early spring. Our review of projected snow persistence (to approximately year 2055) within the Northern and Southern Rocky Mountains indicates that several hundred square kilometers/miles of snow at a depth of 20 inches or more will persist on May 1 at the elevations used by wolverines for denning.

Our snow retention and climate assessments were conducted by the National Oceanic and Atmospheric Administration and the University of Colorado Boulder’s Cooperative Institute for Research in Environmental Sciences. For more information, see page 73 of our species status assessment.

It is also important to note that new wolverine research has provided evidence that snow cover is not as critical as previously thought to denning, with individuals observed outside previously modeled projections of spring snow cover.

Q: What new information have scientists learned about the wolverine since the Service’s February 2013 proposal?

A: Several new wolverine studies have been published in this time, improving our understanding of wolverine biology while also highlighting new insights into the species’ environmental needs. A complete list of citations is available in our species status assessment.

Of particular note, wolverine populations and wolverine dens have been observed outside previously modeled projections of spring snow cover, making the presence of snow for denning not as important as previously thought. In addition, in the species status assessment, we evaluated the potential impacts of legal trapping in southern Canada, an analysis that was not conducted for the 2013 proposal or the 2014 withdrawal. Based on this new analysis, we find that legal trapping does not represent a barrier or stressor to wolverines migrating into the United States at the individual or population level.

Q: What basic elements do wolverines need to survive in the U.S.?

A: Overall, the best available information indicates that the wolverine’s physical and ecological needs in the contiguous United States include:

  • Large territories in relatively inaccessible landscapes; at high elevation (5,906 to 11,483 feet.)
  • Access to a variety of food resources that vary with seasons.
  • Physical and structural features (e.g., talus slopes, rugged terrain) linked to reproductive behavioral patterns.

Q: How can the Service make a determination that the wolverine is not threatened or endangered without accurate population numbers?

A: Actual current population size, growth rate and current population trends are unknown for wolverines in the contiguous United States due to the lack of abundance information and the natural behavior of the species. However, the best available information does not indicate either increasing or declining numbers of the wolverine in North America, including the contiguous United States. Currently, we are seeing expansion of wolverines into historically occupied areas in the contiguous United States. Further, at this time, the best available information does not indicate that the species’ abundance is significantly impacted by the stressors evaluated (singly or cumulatively), and this situation is unlikely to change in the future, supporting current and future resiliency.

Q: Why does the forecast stop at 2055? What is projected to happen to wolverines after then as effects of climate change increase?

A: The future timeframe evaluated in our SSA analysis is approximately 38 to 50 years, which captures consideration of the projected future conditions related to trapping/harvesting, climate change, or other potential cumulative impacts (Service 2018, p. 73) through 2055. We believe this is a reasonable timeframe to consider for our analysis as it also includes the potential for observing these effects over several generations of the wolverine. We cannot speculate beyond the foreseeable future in making listing determinations.

 

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Archives »

March 27, 2018: Collaborative studies conducted on the Wind River Reservation in Wyoming confirm the presence of wolverines there.




October 17, 2016: The U.S. Fish and Wildlife Service (Service) is reopening the public comment period on a proposed rule to list the North American wolverine as threatened under the Endangered Species Act (ESA). 





April 4, 2016: The United States District Court for the District of Montana overturned our decision to withdraw our 2014 proposal to list the wolverine as threatened, remanding the decision back to the Service for further consideration.




August 13, 2014: The Service withdrew a proposal to list the North American wolverine in the contiguous United States as a threatened species under the Endangered Species Act (ESA). The wolverine, a large but elusive member of the weasel family found in the Mountain West, has made a steady recovery in the past half century after hunting, trapping and poisoning nearly extirpated the species from the lower 48 states in the early 1900s.

While it is clear that the climate is warming, after carefully considering the best available science, the Service has determined that the effects of climate change are not likely to place the wolverine in danger of extinction now or in the foreseeable future. As a result, the wolverine does not meet the statutory definition of either a “threatened species” or an “endangered species” and does not warrant protection under the ESA.




April 3 and 4, 2014: The Service convened a panel of scientific experts to provide input on the potential effects of future climate changes on wolverines and their habitat. The panel was organized in response to peer review and state comments we received after publication of the proposed rule to list wolverines. We are posting the results here so that people may provide comments to us about these results prior to the end the of the comment period which closes on May 6th, 2014. Please go to regulations.gov and search for "wolverine" to send us your comments.




February 4, 2014: The U.S. Fish and Wildlife Service published a Federal Register notice that will extend the deadline for our final decision on whether to list the wolverine under the Endangered Species Act. The Act allows for such an extension when there is substantial scientific disagreement regarding the sufficiency or accuracy of the available data relevant to the decision at issue. During the peer review process on our proposed rule to list the wolverine as threatened, we received a variety of opinions from the scientific community concerning the information we used to develop the proposed rules. In response, we will be extending the deadline for the final listing decision by 6 months to further evaluate areas of scientific disagreement and uncertainty as they relate to the wolverine listing decision.

The new deadline for a final rule or withdrawal of the proposal will be August 4, 2014. With the Federal Register notice announcing the 6-month extension we are also reopening the comment period for the wolverine listing until May 6, 2014, details on the kinds of information the Service is seeking is available in the 6-month extension notice.




February 4, 2013: Wolverines were nearly extirpated from the contiguous United States in the early 20th century due to broad-scale predator trapping and poisoning programs.  Since that time they have made a remarkable recovery. Unfortunately, climate warming over the next century is likely to significantly reduce wolverine habitat, to the point where persistence of wolverines in the contiguous United States, without intervention, is in doubt.  Endangered Species Act (ESA) protection and actions by Federal and State agencies in partnership with private landowners and nongovernmental organizations can help protect the wolverine from extinction by increasing its ability to persist in the face of climate change.  

We are, therefore, proposing to protect the North American wolverine as a threatened species under the ESA.  

We are also proposing a special rule that would limit protections of the ESA only to those necessary to address the threats to the species. In the case of the wolverine, human activities in wolverine habitat such as snowmobiling, backcountry skiing, and land management activities like timber harvest and infrastructure development, which do not constitute threats to the species, would not be prohibited or regulated.  However, intentional killing of wolverines would be prohibited.   We are seeking input on whether or not it is appropriate to prohibit incidental take of wolverine in the course of legal trapping activities directed at other species, if states have programs in place to minimize the chances of this occurring. 

A 90-day comment period, beginning February 4, 2013, is being provided to allow the public and stakeholders an opportunity to comment on these proposals.  During that time, we will also seek peer review from qualified members of the scientific community to ensure that our final decision is based on solid science. 

The Service will make a final determination a year from now on whether to add the wolverine to the Federal List of Endangered and Threatened Wildlife based on the best available science. The Service will also decide whether or not it is prudent to designate critical habitat for the wolverine, and whether such a designation would be beneficial to this species given the threat to its habitat is climate change. 

Information on how to submit public comments, including information on the public meetings and hearings that we will hold on the proposed rulemakings is provided below.

Scientific information regarding these proposals will be accepted until May 6, 2013 and can be submitted electronically via the Federal eRulemaking Portal at http://www.regulations.gov. Follow the instructions for submitting comments; or mailed or hand delivered to: Public Comments Processing, Attn: FWS-R6-ES-2012-0107; Division of Policy and Directives Management; U.S. Fish and Wildlife Service; 4401 N. Fairfax Drive, Suite 222; Arlington, VA 22203.

Public Hearings: Three informational sessions and public hearing will be held on this proposed rule.

  • Boise, Idaho:  March 13, 2013 at the Boise Centre on the Grove, 850 West Front Street, Boise, ID 83702.
  • Lakewood, Colorado: March 19, 2013 at the Hampton Inn, 137 Union Boulevard, Lakewood, CO 80228.  
  • Helena, Montana: March 27, 2013 at the Red Lion Colonial Inn, 2301 Colonial Drive, Helena, MT 59601.  

At all three locations the public informational session will run from 2:00 PM to 5:00 PM, followed by public speaker registration at 6:00 PM, and then the public hearing for oral testimony from 7:00 PM to 9:00 PM. 



December 2010: After a thorough review of all the available science, the Fish and Wildlife Service has determined that the contiguous United States population of wolverine should be protected under the Endangered Species Act (ESA). However, the rulemaking to propose ESA protections for the wolverine will be delayed while we work on listing proposals for other species in greater need. The wolverine will be added to the list of candidates for ESA protection. As a candidate species, the wolverine will not receive protection under the ESA; however, we will review its status annually and will continue to work with landowners and partners to implement voluntary conservation measures.

The results of status review indicate that climate warming is the primary threat to wolverine. Our evaluation found that the effects of climate warming are serious but so far have not resulted in any detectable population effects to the species. Because the threat of climate warming is not imminent, we will use our resources to work on listing determinations for species at greater risk of extinction.




April 2010: The U.S. Fish and Wildlife Service has initiated a status review of the wolverine to determine whether the species warrants protection under the Endangered Species Act (ESA).  

The status review will examine potential wolverine populations in California, Colorado, Idaho, Montana, Oregon, Washington and Wyoming.

The Service is seeking the latest scientific and commercial information on the status of the wolverine from the public, government agencies, tribes, industry and the scientific and conservation communities. After gathering and analyzing this information, the Service will determine whether to propose adding the wolverine to the federal list of threatened and endangered species. Information will be received until May 17, 2010.




March 2008: "Not Warranted" Finding - In a finding published March 11, 2008, the U.S. Fish and Wildlife Service determined that the contiguous United States population of wolverine is not a listable entity under the Endangered Species Act and therefore not eligible for protection under the Act.

The Service continues to seek new information regarding the status of the wolverine and continues to support cooperative conservation efforts to benefit the species in its native range.




2007: Status Review - The U.S. Fish and Wildlife Service announced it is initiating a status review of the wolverine (Gulo gulo luscus) to determine whether the species wattants protection under the Endangered Species Act.




October 2003: Wolverine Petition "Not Substantial" - The Biodiversity Legal Foundation and five other organizations petitioned the Service to list the wolverine within the contiguous United States and designate critical habitat.  The Service determined that the petition and other currently available data is insufficient to determine wolverine distribution, habitat requirements, and whether there are threats to the continued existence of the wolverine.


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The mission of the U.S. Fish and Wildlife Service is working with Others to conserve, protect, and enhance fish, wildlife, plants and their habitats for the continuing benefit of the American People.
Last modified: January 06, 2021
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