Nevada Fish & Wildlife Office
Pacific Southwest Region

Desert Tortoise Recovery

DTRO Staff Publications

  • Averill-Murray, R.C., T.E. Christopher, and B.T. Henen. 2018. Reproductive ecology and life history of female Sonoran Desert Tortoises (Gopherus morafkai). Herpetological Monographs 32:34–50. (0.475MB PDF)
  • Campbell, S., E. Zylstra, C. Darst, R. Averill-Murray, and R. Steidl. 2018. A spatially explicit hierarchical model to characterize population viability. Ecological Applications 28:2055–2065. (0.612MB PDF)
  • Allison, L.J., and A.M. Luckie. 2018. Population trends in Mojave Desert Tortoises (Gopherus agassizii). Herpetological Conservation and Biology 13:433-42. (1.3MB PDF)
  • Field K.J., J. Johnson, and N. Lamberski. 2018. Nasal-oral water administration of juvenile Mohave Desert Tortoises. Journal of Fish and Wildlife Management 9:xx-xx. doi:10.3996/042017-JFWM-034. (1.6MB PDF) (1.3MB PDF)
  • Lovich, J.E., R. Averill-Murray, M. Agha, J.R. Ennen, and M. Austin. 2017. Variation in Annual Clutch Phenology of Desert Tortoises (Gopherus morafkai) in the Sonoran Desert of Arizona. Herpetologica 73:313-322.
  • Ennen, J.R., J.E. Lovich, R.C. Averill-Murray, C. Yackulic, M. Agha, C. Loughran, L. Tennant, and B. Sinervo. 2017. The evolution of different maternal investment strategies in two closely related desert vertebrates. Ecology and Evolution doi:10.1002/ece3.2838 (.58 MB PDF)
  • Nafus, M.G., T.C. Esque, R.C. Averill-Murray, K.E. Nussear, and R.R. Swaisgood. 2016. Habitat drives dispersal and survival of translocated juvenile desert tortoises. Journal of Applied Ecology doi:10.1111/1365-2664.12774
  • Germano, J.M., K.J. Field, R.A. Griffiths, S. Clulow, J. Foster, G. Harding, and R.R. Swaisgood. 2015. Mitigation-driven translocations: are we moving wildlife in the right directions? Frontiers in Ecology and the Environment 13:100-105. doi:10.1890/140137
  • Proença, L.M., S. Fowler, S. Kleine, J. Quandt, C.O. Mullen, and S.J. Divers. 2014. Single surgeon coelioscopic orchiectomy of desert tortoises (Gopherus agassizii) for population management. Veterinary Record. doi: 10.1136/vr.102421.
  • Averill-Murray, R.C., and B.E. Hagerty. 2014. Translocation relative to spatial genetic structure of the Mojave Desert Tortoise, Gopherus agassizii. Chelonian Conservation and Biology 13:35-41.
  • Sullivan, B.K., R.C. Averill-Murray, K.O. Sullivan, J.R. Sullivan, E.A. Sullivan, and J.D. Riedle. 2014. Winter activity of the Sonoran Desert Tortoise (Gopherus morafkai (22MB PDF)
  • Allison, L.J., and E.D. McCoy. 2014. North American tortoise abundance. Pages 118-126 in D.C. Rostal, E.D. McCoy, and H.R. Mushinsky (eds.), Biology and Conservation of North American Tortoises. Johns Hopkins Univ. Press.
  • Averill-Murray, R.C., L.J. Allison, and L.L. Smith. 2014. Nesting and reproductive output among North American tortoises. Pages 110-117 in D.C. Rostal, E.D. McCoy, and H.R. Mushinsky (eds.), Biology and Conservation of North American Tortoises. Johns Hopkins Univ. Press.
  • Averill-Murray, R.C., Darst C.R., N. Strout, and M. Wong. 2013. Conserving population linkages for the Mojave Desert Tortoise (Gopherus agassizii). Herpetological Conservation and Biology 8:1-15. (1.3MB PDF)

  • Darst C.R., P.J. Murphy, N.W. Strout, S.P. Campbell, K.J. Field, L. Allison, and R.C. Averill-Murray. 2013. A strategy for prioritizing threats and recovery actions for at-risk species. Environmental Management 51:786-800

  • Zylstra, E.R., R.J. Steidl, C.A. Jones, and R.C. Averill-Murray. 2012. Spatial and temporal variation in survival of a rare reptile: a 22-year study of Sonoran Desert Tortoises. Oecologica 173:107-116.
  • Averill-Murray, R.C., C.R. Darst, K.J. Field, and L.J. Allison. 2012. A new approach to conservation of the Mojave Desert Tortoise. BioScience 62:893-899.
  • Averill-Murray, R.C. 2011. Comment on the conservation status of the desert tortoise(s). Herpetological Review 42:500-501. (PDF 1.2MB)
  • Esque, T.C., K.E. Nussear, K.K. Drake, A.D. Walde, K.H. Berry, R.C. Averill-Murray, A.P. Woodman, W.I. Boarman, P.A. Medica, J. Mack, and J.S. Heaton. 2010. Effects of subsidized predators, resource variability, and human population density on desert tortoise populations in the Mojave Desert, USA. Endangered Species Research 12:167-177.
  • Riedle, J.D., R.C. Averill-Murray, and D.D. Grandmaison. 2010. Seasonal variation in survivorship and mortality of Desert Tortoises in the Sonoran Desert, Arizona. Journal of Herpetology 44:164-167.
  • Reed, J.M., N. Fefferman, and R.C. Averill-Murray. 2009. Vital rate sensitivity analysis as a tool for assessing management actions for the Desert Tortoise. Biological Conservation 142:2710-2717.
  • Riedle, J.D., R.C. Averill-Murray, C.L. Lutz, and D.K. Bolen. 2008. Habitat use by Desert Tortoises (Gopherus agassizii) on alluvial fans in the Sonoran Desert, Arizona. Copeia 2008:414-420. (PDF .554MB)
  • Field, K.J., C.R. Tracy, P.A. Medica, R.W. Marlow, and P.S. Corn. 2007. Return to the wild: translocation as a tool in conservation of the Desert Tortoise (Gopherus agassizii). Biological Conservation 136:232-245.
  • Averill-Murray, R.C., and A. Averill-Murray. 2005. Regional-scale estimation of density and habitat use of the Desert Tortoise (Gopherus agassizii) in Arizona. Journal of Herpetology 39:65-72.

  • Averill-Murray, R.C. 2002. Effects on survival of Desert Tortoises (Gopherus agassizii) urinating during handling. Chelonian Conservation and Biology 4:430-435.
  • Averill-Murray, R.C. 2002. Reproduction of Gopherus agassizii in the Sonoran Desert, Arizona. Chelonian Conservation and Biology 4:295-301.
  • Averill-Murray, R.C., B.E. Martin, S.J. Bailey, and E.B. Wirt. 2002. Activity and behavior of the Sonoran Desert Tortoise in Arizona. Pages 135-158 in T.R. Van Devender (ed.), The Sonoran Desert Tortoise: Natural History, Biology, and Conservation. Univ. Arizona, Tucson.
  • Averill-Murray, R.C., A.P. Woodman, and J.M. Howland. 2002. Population ecology of the Sonoran Desert Tortoise in Arizona. Pages 109-134 in T.R. Van Devender (ed.), The Sonoran Desert Tortoise: Natural History, Biology, and Conservation. Univ. Arizona, Tucson.
  • Swann, D.E., R.C. Averill-Murray, and C.R. Schwalbe. 2002. Distance sampling for Sonoran Desert Tortoises. Journal of Wildlife Management 66:969-975.
  • Van Devender, T.R., Averill-Murray, R.C., T.C. Esque, P.A. Holm, V.M. Dickinson, C.R. Schwalbe, E.B. Wirt, and S.L. Barrett. 2002. Grasses, mallows, desert vine, and more: diet of the Desert Tortoise in Arizona and Sonora. Pages 159-193 in T.R. Van Devender (ed.), The Sonoran Desert Tortoise: Natural History, Biology, and Conservation. Univ. Arizona, Tucson.
  • Johnson, J.D., R.C. Averill-Murray, and J.L. Jarchow. 2001. Captive care of the Desert Tortoise, Gopherus agassizii. Journal of Herpetological Medicine and Surgery 11:8-15. (PDF 2 MB)
  • Averill-Murray, R.C. 1997. Gopherus agassizii (Desert Tortoise). Diet. Herpetological Review 28:87.

  • Averill-Murray, R.C., C.R. Schwalbe, S.J. Bailey, S.P. Cuneo, and S.D. Hart. 1996. Reproduction in a population of the Desert Tortoise, Gopherus agassizii, in the Sonoran Desert. Herpetological Natural History 4:83-88.


    Additional DTRO Staff Publications

  • Averill-Murray, R.C., and A. Averill-Murray. 2017. Notes on activity and habitat use of Pholidobolus vertebralis (Squamata: Gymnophthalmidae) at a cloud forest reserve in Ecuador. Herpetological Review 48:728–730. 

  • Averill-Murray, R.C., and A. Averill-Murray. 2017. Atractus dunni (Sleepy Groundsnake). Maximum size. Herpetological Review 48:665.

  • Averill-Murray, R.C., and A. Averill-Murray. 2017. Urotheca lateristriga (Ribboned Brittle-snake). Maximum size. Herpetological Review 48:687.

  • McKinney, T., D.E. Brown, and L.J. Allison. 2008. Winter precipitation and recruitment of pronghorns in Arizona. Southwestern Naturalist 53:319-325.
  • Retallick, R.W.R., V. Miera, K.L. Richards, K.J. Field, J.P. Collins. 2006. A non-lethal technique for detecting the chytrid fungus Batrachochytrium dendrobatidis on tadpoles. Diseases of Aquatic Organisms 72:77-85.
  • Field, K.J., and D. Groebner. 2005. Rana chiricahuensis (Chiricahua Leopard Frog). Reproduction. Herpetological Review. 36:306-307.
  • Sredl, M.J., and K.J. Field. 2005. Gastrophryne olivacea (Hallowell, 1857 "1856") Western Narrow-mouthed Toad. Pages 503-506 in M. Lannoo (ed), Amphibian Declines: The Conservation Status of United States Species. University of California Press, Berkeley, CA.
  • Goldberg, C.S., K.J. Field, and M.J. Sredl. 2004. Mitochondrial DNA sequences do not support species status of the Ramsey Canyon Leopard Frog (Rana subaquavocalis). Journal of Herpetology. 38:313-319.
  • Allison, L.J., C.E. Paradzick, J.W. Rourke, and T.D. McCarthy. 2003. A characterization of vegetation in nesting and non-nesting plots for Southwestern Willow Flycatchers in Central Arizona. Studies in Avian Biology 26:81-90.
  • Davidson, R.F., and L.J. Allison. 2003. Effects of monogamy and polygyny on reproductive success in Southwestern Willow Flycatchers (Empidonax traillii extimus) in Arizona. Studies in Avian Biology 26:118-124.
  • Field, K.J., T.L. Beatty Sr., and T.L. Beatty Jr. 2003. Rana subaquavocalis (Ramsey Canyon Leopard Frog). Diet. Herpetological Review 34:235.
  • Nussear, K.E., R.E. Espinoza, C.M. Gubbins, K.J. Field, and J.P. Hayes. 1998. Diet quality does not affect resting metabolic rate or body temperatures selected by an herbivorous lizard. Journal of Comparative Physiology B 168:183-189. [see Erratum:1999.169:236]
  • Allison, L.J., P.E. Brunkow, and J.P. Collins. 1994. An exceptional life history trait in tiger salamanders: opportunistic breeding after summer rains. Great Basin Naturalist 54:376-379.
  • Whiting, M.J., J.R. Dixon, and Averill-Murray, R.C. 1993. Spatial distribution of a population of Texas Horned Lizards (Phrynosoma cornutum: Phrynosomatidae) relative to habitat and prey. Southwestern Naturalist 38:150-154.
    Last updated: May 20, 2019