CLASSIFICATION:
Federal Endangered Species (Federal Register
50:4222; January 30, 1985)
CRITICAL HABITAT: Designated in Federal Register 50:4222;
January 30, 1985. (see pdf map)
Critical habitat includes 857 acres in Fresno County. Twenty-three acres are in
a small part of the Mendota Wildlife Management Area, 732 acres comprise the
contiguous Alkali Sink ER, both State-owned and managed, and 102 acres are
in five privately-owned parcels.
DESCRIPTION: The Fresno kangaroo rat (Dipodomys nitratoides exilis) is one of three subspecies of San Joaquin kangaroo
rats, distinguished by being smaller than the Tipton (Dipodomys nitratoides nitratoides) and the short-nosed kangaroo
rat (Dipodomys nitratoides brevinasus), a species
of concern. The Fresno subspecies averages around nine inches in length. San
Joaquin kangaroo rats can be distinguished from other kangaroo rats within their
range by the presence of four toes on the hind foot; other species have five toes.
Kangaroo rats are in the family Heteromyidae. They are not really rats at
all. At least, they are not like common nonnative household rats, which are in
the Muridae family.
Kangaroo rat adaptations for two-footed hopping include elongated hind limbs, a long,
tufted tail for balance, a shortened neck and a large, flattened head. Other
characteristics include large, dorsally placed eyes and small, rounded ears. Fore-limbs are comparatively short with stout claws that facilitate digging burrows. The fur is dark yellowish-buff dorsally and white ventrally. A
white stripe extends across the hips, continuing for the length of the tufted tail.
The base of the tail is circumscribed by white. The top and bottom of the tail
are blackish. Dark whisker patches on each side of the nose are connected by a
black band of fur.
Fresno kangaroo rats eat mostly seeds, with small amounts of green,
herbaceous vegetation and insects supplementing their diet when available.
Most kangaroo rats gather seeds when they are available and cache them for
consumption later. Typically, caches are made in small pits that hold the
contents of the two cheek pouches. Caches are located on the surface of the
soil, and are typically scattered over the home range of the individual. Fresno
kangaroo rats may not cache seeds in their burrows to the same extent as other
kangaroo rats. The soil where they live is damp much of the year, and seeds
would spoil rapidly under such conditions. Therefore, Fresno kangaroo rats may
be obligated to forage on the surface year round.
Little is known about San Joaquin kangaroo rat reproduction in the wild.
Mating appears to begin in the winter. Most females seem to have one litter per
year, although some have two or more. Young are born in burrows.
DISTRIBUTION: The historic range of the Fresno kangaroo rat encompassed an area of grassland
and chenopod scrub communities on the San Joaquin Valley floor, from about the
Merced River, Merced County, on the north, to the northern edge of the marshes
surrounding Tulare Lake, Kings County, on the south, and extending from the
edge of the Valley floor near Livingston, Madera, Fresno, and Selma, westward to
the wetlands of Fresno Slough and the San Joaquin River. Documentation of
historical distribution is scanty. An estimate of the historical range, within the
area as outlined above, is about 888,459 acres. As Fresno kangaroo rats prefer
nearly level, light friable soils, not all of this area would have been habitat
There are no known populations within the historical geographic
range in Merced, Madera and Fresno counties. In spring of 1986 a levee on the
south side of the San Joaquin River broke, flooding the Alkali Sink Ecological
Reserve and other habitat designated as critical habitat. Water approximately 3
feet deep covered most of the area for several days. Trapping between 1988
and 1992 failed to locate any Fresno kangaroo rats on the Alkali Sink Ecological
Reserve. In the autumn of 1992 a single male was captured twice; trapping in
1993, 1994 and 1995 has not yielded additional captures. Trapping at other
sites in Merced, Madera, and Fresno counties between 1988 and 1995 has
failed to locate other extant populations.
In Kings County, two populations of San Joaquin kangaroo rats have been found
on about 371 acres in 1994 and 1995. One site, Lemoore Naval Air Station, is
97 acres. Whether these populations belong to the Fresno or Tipton subspecies
is uncertain, but historically, their ranges were contiguous.
SPECIAL CONSIDERATIONS: Loss of habitat to cultivation, year-round grazing and conversion of land to other
uses, coupled with the resulting fragmentation and isolation of populations
increase the probability of extinction.
Flooding due to levee failure also poses a high risk because of the proximity to
the San Joaquin River. Other potential threats are the indiscriminate use of
rodenticides, competition with Heermann's kangaroo rats and disease and
predation, any of which could extirpate small, isolated populations.
REFERENCES FOR ADDITIONAL INFORMATION: Best, T.L. 1991. Dipodomys nitratoides. Mammal. Species 381:1-7.
Brown, N.L. and D.F. Williams. Endangered Species Recovery
Program. Species account.
Thelander, C. ed. 1994. Life on the edge: a guide to California's endangered
natural resources. BioSystem Books. Santa Cruz, CA. p 76-77.
U.S. Fish and Wildlife Service. 1998.Recovery plan for the upland species of the San
Joaquin Valley, California, September 30. 1998. (pdf format)Portland, OR.
Photo Credit:Moose Peterson©
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